Main investigator and partner
![Nadine Ruppert
Head of the project](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Nadine-Ruppert-2017.jpg)
Head of the project
Malaysia
![Antje Engelhardt
Main collaboration partner and advisor](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Antje-Engelhardt-2015.jpg)
Main collaboration partner and advisor
Germany
![Anja Widdig
Collaboration partner
University Leipzig & Max-Planck Institute for Evolutionary Anthropology](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Anja-Widdig-2017.jpg)
Collaboration partner University
Leipzig & Max-Planck Institute for Evolutionary Anthropology
Germany
![Asyraf Mansor
Co-investigator
School of Biological Sciences, Universiti Sains Malaysia](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Asyraf-Mansor_01.jpg)
Co-investigator
School of Biological Sciences
Universiti Sains Malaysia
Malaysia
![Shahrul Anuar Mohd Sah
Co-researcher
School of Biological Sciences, Universiti Sains Malaysia](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Shahrul-Anuar-Mohd-Sah.jpg)
School of Biological Sciences,
Universiti Sains Malaysia
Malaysia
![](http://macaca-nemestrina.org/wp-content/uploads/2023/02/avatar_neutral.png)
Research collaborator
Central Washington University
USA
![](http://macaca-nemestrina.org/wp-content/uploads/2023/02/avatar_neutral.png)
Research collaborator
University of California Davies
USA
![Malaysian Primatological Society
Local collaborator](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Malaysian-Primatological-society.jpg)
Local collaborator
Malaysia
![](http://macaca-nemestrina.org/wp-content/uploads/2023/02/avatar_neutral.png)
Local collaborator
Malaysia
![](http://macaca-nemestrina.org/wp-content/uploads/2023/02/avatar_neutral.png)
Local collaborator
Malaysia
![Fuze Ecoteer
Local collaborator](http://macaca-nemestrina.org/wp-content/uploads/2023/02/FuzeEcoteerBanner.png)
Malaysia
Students and volunteers
![Michele Olsthoorn
Intern
2017
Netherland](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Michele-Olsthoorn.jpg)
Intern 2017
Netherland
![Megha Majoe
Volunteer
2017
India](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Mega-Majoe.jpg)
Volunteer 2017
India
![Anna Holzner
PhD student
since 2017
Germany](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Anna-Holzner.jpg)
PhD student
since 2017
Germany
![](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Avatar-Macaca-nemestrinat.jpg)
PhD student
2017
Italy
![Emily Dura
Master student
2016 / 2017
USA](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Emily-Dura.jpg)
Master student
2016 / 2017
USA
![Amira Rahman
Research assistant
2015 - 2017
Malaysia](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Amira-Rahman.jpg)
Research assistant
2015 – 2017
Malaysia
![Samantha Gate
BA student
2016
United Kingdom](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Samantha-Gate.jpg)
BA student
2016
United Kingdom
![Bas van Boekholt
Volunteer
2015 / 2016
Netherlands](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Bas-van-Boekholt.jpg)
Volunteer
2015 / 2016
Netherlands
![Simone Luehr
Volunteer
2015
Germany](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Simone-Luehr.jpg)
Volunteer
2015
Germany
![Janine Raethke
Volunteer
2015
Germany](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Janine-Raethke.jpg)
Volunteer
2015
Germany
![Khai Wei See
Volunteer
2015
Malaysia](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Khai-Wei-See.jpg)
Volunteer
2015
Malaysia
![Filip Swat
Volunteer
2015
USA](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Filip-Swat.jpg)
Volunteer
2015
USA
![Laura Morgan
Volunteer
2015
Belgium](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Laura-Morgan.jpg)
Volunteer
2015
Belgium
![Viola Eckerle
Volunteer
2015
Germany](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Viola-Eckerle.jpg)
Volunteer
2015
Germany
![Philipp Hoenle
Volunteer
2014/2015
Germany](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Phil-Hoenle.jpg)
Volunteer
2014/2015
Germanyy
![Eva Tankovic
Volunteer
2014/2015
France](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Eva-Tankovic.jpg)
Volunteer
2014/2015
France
![Léa Radzik
Volunteer
2014/2015
France](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Lea-Radzik-01.jpg)
Volunteer
2014/2015
FranceLéa Radzik Volunteer 2014/2015 France
![Emelie Mayens
Bachelor candidate
2014/2015
Belgium](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Emelie-Mayens.jpg)
Bachelor candidate
2014/2015
Belgium
![Alice Slade
Volunteer
2014/2015
France](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Alice-Slade.jpg)
Volunteer
2014/2015
United Kingdom
![Aurelie Coeut
Volunteer
2014
France](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Aurelie-Coeut.jpg)
Volunteer
2014
France
![Zoe Beaumont
´Volunteer
2014
France](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Zoe-Beaumont.jpg)
´Volunteer
2014
France
![Stephanie Kordon
Volunteer
2014
Austria](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Stephanie-Kordon.jpg)
Volunteer
2014
Austria
![Maximilian Luebben
Volunteer
2014
Germany](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Maximilian-Luebben.jpg)
Volunteer
2014
Germany
![Valentin Erlenbach
Volunteer
2013/2014
Germany](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Valentin-Erlenbach.jpg)
Volunteer
2013/2014
Germany
![Michaela Schneller
Volunteer
2014
Germany](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Michaela-Schneller.jpg)
Volunteer
2014
Germany
![Sandra Schweiger
Volunteer
2013/2014
Germany](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Sandra-Schweiger.jpg)
Volunteer
2013/2014
Germany
![](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Avatar-Macaca-nemestrinat.jpg)
Volunteer
2013
Germany
![Matthias Schuster
Volunteer
2013
Germany](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Anna-Gisbrecht.jpg)
Volunteer
2013
Germany
![Anna Gisbrecht
Master student
2013/2014
Germany](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Carolin-Kade.jpg)
Volunteer
2013
Germany
![Luis Vendrell-Ciurana
Volunteer
2013
Spain](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Luis-Vendrell-Ciurana.jpg)
2013
Spain
![Julia Borstel
Volunteer
2012/2013
Germany](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Julia-Borstel.jpg)
Volunteer
2012/2013
Germany
![Angelika Beck
Master student
2012/2013
Germany](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Angelika-Beck.jpg)
Master student
2012/2013
Germany
![Martin Heinl
Volunteer
2012/2013
Germany](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Martin-Heinl_01.jpg)
Volunteer
2012/2013
Germany
![Benediktus Giyarto (†)
Research manager field station "Tangkoko", Indonesia
2012
Indonesia](http://macaca-nemestrina.org/wp-content/uploads/2023/02/Benediktus-Ugiek-Giyarto-02.jpg)
Research manager field station „Tangkoko“,
2012
Indonesia